Jonathan Sexton has been named at out-half ahead of Ronan O'Gara for Ireland's Test match against Australia in Brisbane on Saturday.
The Leinster No 10 was successful with seven out of eight kicks at goal last week in the narrow loss to the New Zealand Maoris and has been retained at the expense of his Munster rival.
Ulster No 8 Chris Henry will win his first cap as coach Declan Kidney makes four changes to the side which capitulated to the All Blacks.
Paddy Wallace steps in for the injured Gordon D'Arcy and Niall Ronan edges out Leinster rookie teenager Rhys Ruddock, who could also earn his first cap from the bench.
Mick O'Driscoll is named at second row after recovering from a back injury to partner Donncha O'Callaghan.